The Taxonomic Status of Lacerta composita Méhely, 1909 (Squamata: Sauria: Lacertidae)

Igor V. Doronin


The purpose of this work, under Articles, 23.9.2, and 24.2 of the International Code of Zoological Nomenclature, is to conserve the priority of the specific name Lacerta mixta Méhely, 1909. This name is threatened by the never-used-as-valid name Lacerta composita Méhely, 1909. It is proposed that Lacerta mixta, Méhely, 1909 is given precedence as a nomen protectum over Lacerta composita Méhely, 1909, which will be considered as a nomen oblitum.


nomenclature; Georgia; rock lizards; Darevskia mixta; nomen oblitum; nomen protectum

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Boulenger G. A. (1920), Monograph of the Lacertidae. Vol. I, British Museum (Natural History) – Trustees of the British Museum (Nat. Hist.), London.

Darevsky I. S. (1967), Rock Lizards of the Caucasus: Systematics, Ecology and Phylogeny of the Polymorphic Groups of Caucasian Rock Lizards of the Subgenus Archaeolacerta. Nauka, Leningrad [in Russian].

International Code of Zoological Nomenclature (1999), The International Trust for Zoological Nomenclature, London.

Méhely L. (1909), «Materialien zu einer Systematik und Phylogenie der muralis-ähnlichen Lacerten», Ann. Mus. National. Hung., 7, 409 – 621, pls. X – XXV.

Mertens R. (1967), «Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen», Senckenberg. Biol., 48(A), 1 – 106.

Nikolsky A. M. (1913), Herpetologia Caucasica, Publications of the Caucasus Museum, Tiflis [in Russian].

Nikolsky A. M. (1915), Faunae de la Russie et des pays limitrophes fondée principalement sur les collections du Musée Zoologique de l’Académie Impériale des Sciences de Petrograd. Reptiles (Reptilia). Vol. I. Chelonia et Sauria, Imperial Academy of Sciences, Petrograd [in Russian].

Radde G. I. (1899), [Die Sammlungen des kaukasischen Museums ] The collections of the Caucasian Museum processed together with scientific experts and published by Doctor G. I. Radde. Vol. I. Zoology, Tipografiya Kantselyarii Glavnonachal’stvuyushchego grazhdanskoy uchast’yu na Kavkaze, Tiflis [in Russian].

Ananjeva N. B., Orlov N. L., Khalikov R. G., Darevsky I. S., Ryabov S. A., and Barabanov A. V. (2006), The Reptiles of Northern Eurasia. Taxonomic Diversity, Distribution, Conservation Status. Pensoft Series Faunistica. Vol. 47, Pensoft, Sofia.

Arakelyan M. S. (2012), «Sympatric speciation processes in populations of bisexual rock lizards of the genus Darevskia», Biol. J. Armenia, 64(3), 10 – 15.

Arnold E. N., Arribas O., and Carranza S. (2007) «Systematics of the Palaearctic and Oriental lizard tribe Lacertini (Squamata: Lacertidae: Lacertinae), with descriptions of eight new genera», Zootaxa, 1430, 1 – 86.

Bischoff W. and Schmidtler J. F. (2014), «Von Pallas bis Darevsky: Die Erforschung der Herpetofauna des Kaukasus am Beispiel der Echsen», Sekretär, 14(2), 23 – 58.

Brown R. P. (2005), «Large subunit mitochondria rRNA secondary structures and site-specific rate variation in two lizard lineages», J. Mol. Evol., 60(1), 45 – 56.

Freitas S., Rocha S., Campos J., Ahmadzadeh F., Corti C., Sillero N., Ilgaz З., Kumlutas Y., Arakelyan M., Harris D. J., and Carretero M. A. (2016), «Parthenogenesis through the ice ages: A biogeographic analysis of Caucasian rock lizards (genus Darevskia)», Mol. Phylogen. Evol., 102, 117 – 127.

Freitas S., Vavakou A., Arakelyan M., Drovetski S. V., Crnobrnja-Isailovic J., Kidov A. A., Cogalniceanu D., Corti C., Lymberakis P., Harris D. J., and Carretero M. A. (2016), «Cryptic diversity and unexpected evolutionary patterns in the meadow lizard, Darevskia praticola (Eversmann, 1834)», Syst. Biodiv., 14(2), 184 – 197.

Gabelaia M., Adriaens D., and Tarkhnishvili D. (2017), «Phylogenetic signals in scale shape in Caucasian rock lizards (Darevskia species)», Zool. Anz., 268, 32 – 40.

Gabelaia M., Tarkhnishvili D., and Murtskhvaladze M. (2015), «Phylogeography and morphological variation in a narrowly distributed Caucasian rock lizard, Darevskia mixta», Amphibia–Reptilia, 36(1), 45 – 54.

Girnyk A. E., Vergun A. A., Omelchenko A. V., Petrosyan V. G., Korchagin V. I., and Ryskov A. P. (2017), «Molecular and genetic characterization of the allelic variants of Du215, Du281, Du323, and Du47G microsatellite loci in parthenogenetic lizard Darevskia armeniaca (Lacertidae)», Russ. J. Gen., 53(4), 472 – 482.

Grechko V. V., Ciobanu D. G., Darevsky I. S., Kosushkin S. A., and Kramerov D. A. (2006), «Molecular evolution of satellite DNA repeats and speciation of lizards of the genus Darevskia (Sauria: Lacertidae)», Genome, 49(10), 1297 – 1307.

Jancúchová-Lásková J., Landová E., and Frynta D. (2015), «Are genetically distinct lizard species able to hybridize? A review», Curr. Zool., 61(1), 155 – 180.

Kosushkin S. A. and Grechko V. V. (2013), «Molecular genetic relationships and some issues of systematics of rock lizards of the genus Darevskia (Squamata: Lacertidae) based on locus analysis on SINE-type repeats (Squam1)», Russ. J. Gen., 49(8), 857 – 869.

Malysheva D. N., Vergun A. A., Martirosyan I. A., Tokarskaya O. N., and Ryskov A. P. (2008), «Molecular and genetic characterization of allelic variants of microsatellite loci in parthenogenetic Caucasian rock lizards Darevskia armeniaca (Lacertidae)», Mol. Gen. Microbiol. Virol., 23(4), 195 – 201.

Panner T. (2014), «Eine herpetologische Reise nach Georgien», Die Eidechse, 25(3), 65 – 75.

Pérez i de Lanuza G. and Font E. (2016), «The evolution of colour pattern complexity: selection for consciousness favours contrasting within-body colour combinations in lizards», J. Evol. Biol., 29(5), 942 – 951.

Sabath N., Itescu Y., Feldman A., Meiri S., Mayrose I., and Valenzuela N. (2016), «Sex determination, longevity and the birth and death of reptilian species», Ecol. Evol., 6(15), 5207 – 5220.

Shacham B., Federman R., Lahav-Ginott S., and Werner Y. L. (2015), «The northward extension of reptiles in the Palearctic, with the Jordan Valley (Israel) as a model: snakes outrace lizards», J. Nat. Hist., 50(15 – 16), 1017 – 1033.

Sindaco R. and Jeremčenko V. K. (2008), The Reptiles of the Western Palearctic. 1. Annotated Checklist and Distributional atlas of the turtles, crocodiles, amphisbaenians and lizards of Europe, North Africa, Middle East and Central Asia, Edizioni Belvedere, Latina (Italy).

Tarkhnishvili D. (2012), «Evolutionary history, habitats, diversification, and speciation in Caucasian rock lizards», Adv. Zool. Res., 2, 79 – 120.

Tarkhnishvili D., Gabelaia M., Mumladze L., and Murtskhvaladze M. (2016), «Mitochondrial phylogeny of the Darevskia saxicola complex: two highly deviant evolutionary lineages from the easternmost part of the range», Herpetol. J., 26(2), 175 – 182.

Tarkhnishvili D., Gavashelishvili A., Avaliani A., Murtskhvaladze M., and Mumladze L. (2010), «Unisexual rock lizard might be outcompeting its bisexual progenitors in the Caucasus», Biol. J. Linn. Soc., 101, 447 – 460.

Tarkhnishvili D., Murtskhvaladze M., and Anderson C. L. (2017), «Coincidence of genotypes at two loci in two parthenogenetic rock lizards. How backcrosses might trigger adaptive speciation», Biol. J. Linn. Soc., 121(2), 365 – 378.

Tuniyev B. S., Lotiev K. J., Tuniyev S. B., Gabaev V. N., and Kidov A. A. (2017), «Amphibians and reptiles of South Ossetia», Nat. Conserv. Res., 2(2), 1 – 23 [in Russian with English summary].

Tuniyev B. S., Orlov N. L., Ananjeva N. B., and Agasyan A. L. (2009), Snakes of the Caucasus: taxonomic diversity, distribution, conservation, KMK Scientific Press, St. Petersburg – Moscow [in Russian].

Tuniyev B. S., Tuniyev S. B., Avcı A., and Ilgaz Ç. (2014), «Herpetological studies in Eastern and North-Eastern Turkey», Curr. Stud. Herpetol., 14(1/2), 44 – 53 [in Russian with English summary].



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