Ecological Observations on the Ezo Brown Frog (Rana pirica Matsui, 1991) on Hokkaido Island, Japan

Sergius L. Kuzmin, Takanori Sato, Shigehiro Nakabayashi, Irina V. Maslova, Nobuyuki Narumi


Observations on reproductive ecology in Rana pirica were conducted in several sites of Hokkaido Island, Japan. Data on adult sizes, phenology, migrations, habitat use, temperatures, and spatial distribution of clutches are discussed in comparison with those on taxonomically related continental species, Rana dybowskii from Russia.


Rana pirica; reproduction; migration; clutch; habitat

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Kuzmin S. L. and Maslova I. V. (2003), The Amphibians of the Russian Far East, Pensoft, Sofia – Moscow.

Maeda N. and Matsui M. (1990), Frogs and Toads of Japan, Bun-Ichi Sogo Shuppan Co., Ltd., Tokyo.

Matsui M. (1991), «Original description of the brown frog from Hokkaido, Japan», Jap. J. Herpetol., 14, 63 – 78.



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